By the wolves in the Western Alps imposed a significant burden on traditional sheep and goat husbandry with a loss of over 5, animals in Northern elephant seal M. The two are however mutually intelligible , as North American wolves have been recorded to respond to European-style howls made by biologists. Also, as wolf and dog mating seasons do not fully coincide, the likelihood of wild wolves and dogs mating and producing surviving offspring is small. Despite periods of intense hunting during the eighteenth century, wolves never disappeared in the western Balkans, from Albania to the former Yugoslavia. In , wolves began recolonising central Sweden after a twelve-year absence, and have since expanded into southern Norway. The population increased again by to about 75,, with 32, being killed in
Its fur is very soft and coarsens with age. The cub begins to crawl at 75 to 80 days;  mothers play with their cubs by rolling and wrestling with them. The cubs can eat small quantities of bamboo after six months,  though mother's milk remains the primary food source for most of the first year. The interval between births in the wild is generally two years. In July , Chinese scientists confirmed the birth of the first cub to be successfully conceived through artificial insemination using frozen sperm.
Attempts have also been made to reproduce giant pandas by interspecific pregnancy by implanting cloned panda embryos into the uterus of an animal of another species. This has resulted in panda fetuses, but no live births. In the past, pandas were thought to be rare and noble creatures — the Empress Dowager Bo was buried with a panda skull in her vault. The grandson of Emperor Taizong of Tang is said to have given Japan two pandas and a sheet of panda skin as a sign of goodwill.
Unlike many other animals in Ancient China , pandas were rarely thought to have medical uses. The few known uses include the Sichuan tribal peoples' use of panda urine to melt accidentally swallowed needles, and the use of panda pelts to control menses as described in the Qin Dynasty encyclopedia Erya.
Zouyu is a legendary "righteous" animal, which, similarly to a qilin , only appears during the rule of a benevolent and sincere monarch. It is said to be fierce as a tiger , but gentle and strictly vegetarian, and described in some books as a white tiger with black spots.
Puzzled about the real zoological identity of the creature captured during the Yongle era, J. Duyvendak exclaims, "Can it possibly have been a Pandah? The comparative obscurity of the giant panda throughout most of China's history is illustrated by the fact that, despite there being a number of depictions of bears in Chinese art starting from its most ancient times, and the bamboo being one of the favorite subjects for Chinese painters, there are no known preth-century artistic representations of giant pandas.
The West first learned of the giant panda on 11 March , when the French missionary Armand David  received a skin from a hunter. The first Westerner known to have seen a living giant panda is the German zoologist Hugo Weigold , who purchased a cub in Kermit and Theodore Roosevelt, Jr.
In , Ruth Harkness became the first Westerner to bring back a live giant panda, a cub named Su Lin  which went to live at the Brookfield Zoo in Chicago. In , five giant pandas were sent to London. Gifts of giant pandas to American and Japanese zoos formed an important part of the diplomacy of the People's Republic of China PRC in the s, as it marked some of the first cultural exchanges between the PRC and the West.
This practice has been termed "panda diplomacy". By , however, pandas were no longer given as gifts. Since , because of a WWF lawsuit , the United States Fish and Wildlife Service only allows a US zoo to import a panda if the zoo can ensure the PRC will channel more than half of its loan fee into conservation efforts for the giant panda and its habitat. The issue became embroiled in cross-Strait relations — both over the underlying symbolism, and over technical issues such as whether the transfer would be considered "domestic" or "international", or whether any true conservation purpose would be served by the exchange.
However, when Ma Ying-jeou assumed the presidency in , the offer was accepted, and the pandas arrived in December of that year. Microbes in panda waste are being investigated for their use in creating biofuels from bamboo and other plant materials. The giant panda is a vulnerable species , threatened by continued habitat loss and habitat fragmentation ,  and by a very low birthrate, both in the wild and in captivity. The giant panda has been a target of poaching by locals since ancient times and by foreigners since it was introduced to the West.
Starting in the s, foreigners were unable to poach giant pandas in China because of the Second Sino-Japanese War and the Chinese Civil War , but pandas remained a source of soft furs for the locals. The population boom in China after created stress on the pandas' habitat, and the subsequent famines led to the increased hunting of wildlife, including pandas. During the Cultural Revolution , all studies and conservation activities on the pandas were stopped.
After the Chinese economic reform , demand for panda skins from Hong Kong and Japan led to illegal poaching for the black market , acts generally ignored by the local officials at the time. Though the Wolong National Nature Reserve was set up by the PRC government in to save the declining panda population, few advances in the conservation of pandas were made, owing to inexperience and insufficient knowledge of ecology. Many believed the best way to save the pandas was to cage them.
As a result, pandas were caged at any sign of decline, and suffered from terrible conditions. Because of pollution and destruction of their natural habitat, along with segregation caused by caging, reproduction of wild pandas was severely limited. In the s, however, several laws including gun control and the removal of resident humans from the reserves helped their chances of survival. With these renewed efforts and improved conservation methods, wild pandas have started to increase in numbers in some areas, though they still are classified as a rare species.
In , scientists reported that the number of pandas living in the wild may have been underestimated at about 1, Previous population surveys had used conventional methods to estimate the size of the wild panda population, but using a new method that analyzes DNA from panda droppings , scientists believe the wild population may be as large as 3, Furthermore, in response to this reclassification, the State Forestry Administration of China announced that they would not accordingly lower the conservation level for panda, and would instead reinforce the conservation efforts.
The giant panda is among the world's most adored and protected rare animals, and is one of the few in the world whose natural inhabitant status was able to gain a UNESCO World Heritage Site designation. The Sichuan Giant Panda Sanctuaries , located in the southwest province of Sichuan and covering seven natural reserves, were inscribed onto the World Heritage List in Not all conservationists agree that the money spent on conserving pandas is well spent.
Chris Packham has argued that the breeding of pandas in captivity is "pointless" because "there is not enough habitat left to sustain them". In , Earthwatch Institute , a global nonprofit that teams volunteers with scientists to conduct important environmental research, launched a program called "On the Trail of Giant Panda".
This program, based in the Wolong National Nature Reserve, allows volunteers to work up close with pandas cared for in captivity, and help them adapt to life in the wild, so that they may breed, and live longer and healthier lives. Pandas have been kept in zoos as early as the Western Han Dynasty in China, where the writer Sima Xiangru noted that the panda was the most treasured animal in the emperor's garden of exotic animals in the capital Chang'an present Xi'an.
Not until the s were pandas again recorded to have been exhibited in China's zoos. Chi Chi at the London Zoo became very popular. This influenced the World Wildlife Fund to use a panda as its symbol. A New York Times article  outlined the economics of keeping pandas, which costs five times more than that of the next most expensive animal, an elephant. San Diego's contract with China was to expire in , but got a five-year extension at about half of the previous yearly cost.
Cryptozoologists use Giant Pandas as an example of an animal recently discovered by science. Skeptical cryptozoologist Joe Nickell , notes that since Giant Pandas were known to local people, they qualify as cryptids. Also, fossil evidence shows that pandas were once widespread, including the two million year old skull of Ailuropoda microta .
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Elsewhere, it can be hunted year-round by Bedouins. Little is known of current wolf populations in Iran, which once occurred throughout the country in low densities during the mids. The northern regions of Afghanistan and Pakistan are important strongholds for the wolf. Overall, India supports about , wolves, scattered among several remnant populations. Although protected since , Indian wolves are classed as endangered, with many populations lingering in low numbers or living in areas increasingly used by humans.
Although present in Nepal and Bhutan , there is no information of wolves occurring there. Wolf populations throughout Northern and Central Asia are largely unknown, but are estimated in the hundreds of thousands based on annual harvests. Since the fall of the Soviet Union , continent-wide extermination of wolves has ceased, and wolf populations have increased to about 25,—30, animals throughout the former Soviet Union.
In China and Mongolia , wolves are only protected in reserves. Mongolian populations have been estimated at 10,—30,, while the status of wolves in China is more fragmentary. The north has a declining population of an estimated wolves, while Xinjiang and Tibet hold about 10, and 2, respectively.
It exists in southern China, which refutes claims made by some researchers in the Western world that the wolf had never existed in southern China. It occurred all over the mainland, save for the southeastern United States, California west of the Sierra Nevada , and the tropical and subtropical areas of Mexico.
Large continental islands occupied by wolves included Newfoundland , Vancouver Island , southeastern Alaskan islands, and throughout the Arctic Archipelago and Greenland. In his November 6, letter to the French Minister of the Marine, Louis Denys de La Ronde reported that the island was home to wolves "of a prodigious size", and sent a wolf pelt back to France to substantiate his claim. As the island was cleared for settlement, the gray wolf population may have been extirpated, or relocated to the mainland across the winter ice: The decline of North American wolf populations coincided with increasing human populations and the expansion of agriculture.
By the start of the 20th century, the species had almost disappeared from the eastern USA, excepting some areas of the Appalachians and the northwestern Great Lakes Region. In Canada, the gray wolf was extirpated in New Brunswick and Nova Scotia between and , and in Newfoundland around It vanished from the southern regions of Quebec and Ontario between and The gray wolf's decline in the prairies began with the extermination of the American bison and other ungulates in the s—70s.
From —, the gray wolf was virtually eliminated from the western USA and adjoining parts of Canada, because of intensive predator control programs aimed at eradicating the species. The gray wolf was exterminated by federal and state governments from all of the USA by , except in Alaska and northern Minnesota. The decline in North American wolf populations was reversed from the s to the early s, particularly in southwestern Canada, because of expanding ungulate populations resulting from improved regulation of big game hunting.
This increase triggered a resumption of wolf control in western and northern Canada. Thousands of wolves were killed from the early s to the early s, mostly by poisoning.
This campaign was halted and wolf populations increased again by the mids. The species' modern range in North America is mostly confined to Alaska and Canada, with populations also occurring in northern Minnesota, northern Wisconsin and Michigan 's Upper Peninsula , and small portions of Washington , Idaho , northern Oregon , and Montana. A functional wolf population should exist in California by according to estimates by state wildlife officials.
In addition, the Mexican wolf Canis lupus baileyi was reintroduced to Arizona and New Mexico in Canada is home to about 52,—60, wolves, whose legal status varies according to province and territory.
First Nations residents may hunt wolves without restriction, and some provinces require licenses for residents to hunt wolves while others do not.
In Alberta , wolves on private land may be baited and hunted by the landowner without requiring a license, and in some areas, wolf hunting bounty programs exist. In Alaska, the gray wolf population is estimated at 6,—7,, and can be legally harvested during hunting and trapping seasons, with bag limits and other restrictions. As of , there are wolves in 28 packs in Yellowstone, and wolves in 25 packs in Idaho.
Reintroduced Mexican wolves in Arizona and New Mexico are protected under the ESA and, as of late , number 28 individuals in eight packs. Viral diseases carried by wolves include rabies , canine distemper , canine parvovirus , infectious canine hepatitis , papillomatosis , and canine coronavirus. Infected wolves do not show any fear of humans, with most documented wolf attacks on people being attributed to rabid animals.
Although canine distemper is lethal in dogs, it has not been recorded to kill wolves, except in Canada and Alaska. The canine parvovirus, which causes death by dehydration , electrolyte imbalance , and endotoxic shock or sepsis , is largely survivable in wolves, but can be lethal to pups.
Wolves may catch infectious canine hepatitis from dogs, though there are no records of wolves dying from it. Papillomatosis has been recorded only once in wolves, and likely doesn't cause serious illness or death, though it may alter feeding behaviors. The canine coronavirus has been recorded in Alaskan wolves, with infections being most prevalent in winter months. Bacterial diseases carried by wolves include brucellosis , lyme disease , leptospirosis , tularemia , bovine tuberculosis ,  listeriosis , anthrax and foot and mouth disease.
While adult wolves tend not to show any clinical signs, it can severely weaken the pups of infected females. Although lyme disease can debilitate individual wolves, it does not appear to have any significant effect on wolf populations. Leptospirosis can be contracted through contact with infected prey or urine, and can cause fever , anorexia , vomiting, anemia , hematuria , icterus , and death.
Wolves living near farms are more vulnerable to the disease than those living in the wilderness, probably because of prolonged contact with infected domestic animal waste.
Wolves may catch tularemia from lagomorph prey, though its effect on wolves is unknown. Although bovine tuberculosis is not considered a major threat to wolves, it has been recorded to have once killed two wolf pups in Canada.
Wolves carry ectoparasites and endoparasites , with wolves in the former Soviet Union having been recorded to carry at least 50 species. In areas where wolves inhabit pastoral areas, the parasites can be spread to livestock. Wolves are often infested with a variety of arthropod exoparasites, including fleas , ticks , lice , and mites.
The most harmful to wolves, particularly pups, is Sarcoptes scabiei or mange mite ,  though they rarely develop full blown mange , unlike foxes. Ticks of the genus Ixodes can infect wolves with Lyme disease and Rocky Mountain spotted fever.
Other ectoparasites include biting lice, sucking lice and the fleas Pulex irritans and Ctenocephalides canis. Endoparasites known to infect wolves include protozoans and helminths flukes , tapeworms , roundworms and thorny-headed worms.
Of 30, protozoan species, only a few have been recorded to infect wolves: Isospora , Toxoplasma , Sarcocystis , Babesia , and Giardia. Upon reaching maturity, Alaria migrates to the wolf's intestine, but harms it little. Metorchis conjunctus , which enters wolves through eating fish, infects the wolf's liver or gall bladder, causing liver disease , inflammation of the pancreas, and emaciation.
Most other fluke species reside in the wolf's intestine, though Paragonimus westermani lives in the lungs. Tapeworms are commonly found in wolves, as their primary hosts are ungulates, small mammals, and fish, which wolves feed upon. Tapeworms generally cause little harm in wolves, though this depends on the number and size of the parasites, and the sensitivity of the host.
Symptoms often include constipation , toxic and allergic reactions , irritation of the intestinal mucosa , and malnutrition. Infections by the tapeworm Echinococcus granulosus in ungulate populations tend to increase in areas with high wolf densities, as wolves can shed Echinoccocus eggs in their feces onto grazing areas. Wolves can carry over 30 roundworm species, though most roundworm infections appear benign, depending on the number of worms and the age of the host.
Ancylostoma caninum attaches itself on the intestinal wall to feed on the host's blood, and can cause hyperchromic anemia , emaciation, diarrhea , and possibly death.
Toxocara canis , a hookworm known to infect wolf pups in utero, can cause intestinal irritation, bloating, vomiting, and diarrhea. Wolves can tolerate low levels of Dirofilaria immitis for many years without showing any ill effects, though high levels can kill wolves through cardiac enlargement and congestive hepatopathy. Wolves probably become infected with Trichinella spiralis by eating infected ungulates.
Nicolla skrjabini , Macrocantorhynchus catulinus , and Moniliformis moniliformis. Human presence appears to stress wolves, as seen by increased cortisol levels in instances such as snowmobiling near their territory. Old English literature contains several instances of Anglo-Saxon kings and warriors taking on wulf as a prefix or suffix in their names. Wolf-related names were also common among pre-Christian Germanic warriors: Ancient Greek literature is similar: Autolycus "the wolf itself" , Lycurgus "wolf-work".
The Latin for "female prostitute" is lupa , and the most famous brothel in Pompeii was the Lupanar. The wolf is a common motif in the foundational mythologies and cosmologies of peoples throughout Eurasia and North America corresponding to the historical extent of the habitat of the gray wolf. The obvious attribute of the wolf is its nature of a predator , and correspondingly it is strongly associated with danger and destruction, making it the symbol of the warrior on one hand, and that of the devil on the other.
The modern trope of the Big Bad Wolf is a development of this. The wolf holds great importance in the cultures and religions of the nomadic peoples, both of the Eurasian steppe and of the North American Plains. In many cultures, the identification of the warrior with the wolf totemism gave rise to the notion of Lycanthropy , the mythical or ritual identification of man and wolf.
Aesop featured wolves in several of his fables , playing on the concerns of Ancient Greece 's settled, sheep-herding world. His most famous is the fable of The Boy Who Cried Wolf , which is directed at those who knowingly raise false alarms, and from which the idiomatic phrase " to cry wolf " is derived.
Some of his other fables concentrate on maintaining the trust between shepherds and guard dogs in their vigilance against wolves, as well as anxieties over the close relationship between wolves and dogs. Although Aesop used wolves to warn, criticize and moralize about human behavior, his portrayals added to the wolf's image as a deceitful and dangerous animal. Although portrayed as loyal, honest and moral, Isengrim is forever the victim of Reynard's wit and cruelty, often dying at the end of each story.
The tale of Little Red Riding Hood , first written in by Charles Perrault , is largely considered to have had more influence than any other source of literature in forging the wolf's negative reputation in the western world.
The wolf in this story is portrayed as a potential rapist , capable of imitating human speech. Tolstoy's War and Peace and Chekhov's Peasants both feature scenes in which wolves are hunted with hounds and borzois. Although credited with having changed popular perceptions on wolves by portraying them as loving, cooperative and noble, it has been criticized for its idealization of wolves and its factual inaccuracies.
The wolf is a frequent charge in English armory. It is illustrated as a supporter on the shields of Lord Welby , Rendel , and Viscount Wolseley , and can be found on the coat of arms of Lovett and the vast majority of the Wilsons and Lows. The demi-wolf is a common crest , appearing in the arms and crests of members of many families, including that of the Wolfes , whose crest depicts a demi-wolf holding a crown in its paws, in reference to the assistance the family gave to Charles II during the battle of Worcester.
Wolf heads are common in Scottish heraldry , particularly in the coats of Clan Robertson and Skene. The wolf is the most common animal in Spanish heraldry , and is often depicted as carrying a lamb in its mouth, or across its back. It is the unofficial symbol of the spetsnaz , and serves as the logo of the Turkish Grey Wolves.
Livestock depredation has been one of the primary reasons for hunting wolves, and can pose a severe problem for wolf conservation: Being the most abundant carnivores, feral and free-ranging dogs have the greatest potential to compete with wolves.
A review of the studies in the competitive effects of dogs on sympatric carnivores did not mention any research on competition between dogs and wolves.
Wolves kill dogs on occasion, with some wolf populations relying on dogs as an important food source. Wolves may display unusually bold behavior when attacking dogs accompanied by people, sometimes ignoring nearby humans.
Large hunting dogs such as Swedish elkhounds are more likely to survive wolf attacks because of their better ability to defend themselves.
Although the numbers of dogs killed each year are relatively low, it induces a fear of wolves entering villages and farmyards to take dogs. In many cultures, there are strong social and emotional bonds between humans and their dogs that can be seen as family members or working team members. The loss of a dog can lead to strong emotional responses with demands for more liberal wolf hunting regulations.
Dogs that are employed to guard sheep help to mitigate human—wolf conflicts, and are often proposed as one of the non-lethal tools in the conservation of wolves.
The fear of wolves has been pervasive in many societies, though humans are not part of the wolf's natural prey. Predatory attacks attacks by wolves treating humans as food may be preceded by a long period of habituation , in which wolves gradually lose their fear of humans. The victims are repeatedly bitten on the head and face, and are then dragged off and consumed, unless the wolves are driven off. Such attacks typically occur only locally, and do not stop until the wolves involved are eliminated.
Predatory attacks can occur at any time of the year, with a peak in the June—August period, when the chances of people entering forested areas for livestock grazing or berry and mushroom picking increase,  though cases of non-rabid wolf attacks in winter have been recorded in Belarus , Kirov and Irkutsk oblasts, Karelia and Ukraine.
The majority of victims of predatory wolf attacks are children under the age of 18 and, in the rare cases where adults are killed, the victims are almost always women. Cases of rabid wolves are low when compared to other species, as wolves do not serve as primary reservoirs of the disease, but can be infected by animals such as dogs, jackals and foxes.
Incidents of rabies in wolves are very rare in North America, though numerous in the eastern Mediterranean , Middle East and Central Asia. Wolves apparently develop the "furious" phase of rabies to a very high degree which, coupled with their size and strength, makes rabid wolves perhaps the most dangerous of rabid animals,  with bites from rabid wolves being 15 times more dangerous than those of rabid dogs.
Most rabid wolf attacks occur in the spring and autumn periods. Unlike with predatory attacks, the victims of rabid wolves are not eaten, and the attacks generally only occur on a single day. The victims are chosen at random, though the majority of cases involve adult men. In the half-century up to , there were eight fatal attacks in Europe and Russia, and more than in south Asia. Wolves are difficult to hunt because of their elusiveness, sharp senses, high endurance, and ability to quickly incapacitate and kill hunting dogs.
This method relies heavily on the wolf's fear of human scents, though it can lose its effectiveness when wolves become accustomed to the smell. Wild wolves are sometimes kept as exotic pets and, in some rarer occasions, as working animals. Although closely related to domesticated dogs , wolves do not show the same tractability as dogs in living alongside humans, and generally, much more work is required in order to obtain the same amount of reliability.
From Wikipedia, the free encyclopedia. Redirected from Gray wolf. This article is about the gray wolf, canis lupus , and its subspecies. For other species of wolf and other uses, see Wolf disambiguation. For other uses, see Grey Wolf disambiguation. Middle Pleistocene — present , years BP . Linnaeus , . Subspecies of Canis lupus.
Evolution of the wolf. List of gray wolf populations by country. Wolves in folklore, religion and mythology. Wolf attacks on humans and List of wolf attacks. Wolf hunting and Wolf hunting with dogs. Human uses of hunted wolves. Wolves as pets and working animals.
However, neither dogs nor dingoes are referred to as gray wolves. Throughout this article, the term "wolf" will be used to refer collectively to naturally occurring subspecies, especially the nominate subspecies, Canis lupus lupus. This terminology was first used in by Rudolf Schenkel of the University of Basel , who based his findings on researching the behavior of captive gray wolves. This view on gray wolf pack dynamics was later popularized by L. David Mech in his book The Wolf. He formally disavowed this terminology in , explaining that it was heavily based on the behavior of captive packs consisting of unrelated individuals, an error reflecting the once prevailing view that wild pack formation occurred in winter among independent gray wolves.
Later research on wild gray wolves revealed that the pack is usually a family consisting of a breeding pair and its offspring of the previous 1—3 years.
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African palm civet N. Angolan slender mongoose G. Ethiopian dwarf mongoose H. Family Viverridae includes Civets. Small-toothed palm civet A. Sulawesi palm civet M. Masked palm civet P. Golden wet-zone palm civet P. Owston's palm civet C. Hose's palm civet D. Banded palm civet H. Malabar large-spotted civet V. Small Indian civet V.
Broad-striped Malagasy mongoose G. American black bear U. Molina's hog-nosed skunk C. Sunda stink badger M. Southern spotted skunk S. Eastern lowland olingo B. Western mountain coati N. South American fur seal A. Northern fur seal C.
Steller sea lion E. Australian sea lion N. South American sea lion O. New Zealand sea lion P. California sea lion Z. Northern elephant seal M. Mediterranean monk seal M. Family Canidae includes dogs. African wild dog L.